American microbiologist and biophysicist Carl Richard Woese proposed three kingdom classification system in 1990. This classification system divides the life forms into three domains and six kingdoms.
The three domains are archaea, bacteria, eukaryote, and six kingdoms are Archaebacteria (ancient bacteria), Eubacteria (true bacteria), Protista, Fungi, Plantae, Animalia.
Woese classified them based on their differences in the 16S ribosomal RNA (rRNA) structure. 16S rRNA can be used for comparative analysis between prokaryotic and eukaryotic species.
Carl Woese used the rRNA as an “Evolutionary Chronometer” – an evolutionary time clock.
The
Archaea (Archaebacteria)
- Archaea are ancient bacteria - believed to have evolved just after the evolution of first life on earth.
- Archaea are prokaryotic cells. The cell walls of Archaea contain no peptidoglycan, hence Archaea are not sensitive to some antibiotics that affect the Bacteria.
- Archaea have membranes composed of branched hydrocarbon chains (many also containing rings within the hydrocarbon chains) attached to glycerol by ether linkages.
- The ether-containing linkages in the Archaea membranes is more stable than the ester-containing linkages in the Eubacteria and are better able to withstand higher temperatures and stronger acid concentrations.
- Archaea often live in extreme environments and include methanogens, extreme halophiles, and hyperthermophiles.
- Archaea contain rRNA that is unique to the Archaea, distinctly different from the rRNA of Bacteria and Eukarya.
Hydrothermal vents on the ocean floor, where the surrounding water can reach over 300° Celsius, are home for some archaeal species.
The Bacteria (Eubacteria)
Bacteria
(also known as eubacteria or "true bacteria") are prokaryotic cells
that are common in human daily life.
Eubacteria can be found almost everywhere and serve as antibiotic producers and food digesters, pathogens etc.
Bacteria are prokaryotic cells. They have membranes composed of unbranched fatty acid chains attached to glycerol by ester linkages.
The cell walls of Bacteria contain peptidoglycan. Bacteria are sensitive to antibacterial antibiotics.
Bacteria
contain rRNA that is unique.
Bacteria
include mycoplasmas, cyanobacteria, Gram-positive bacteria, and Gram-negative
bacteria.
Thus, in short, Archaebacteria are called ancient bacteria
whereas the eubacteria are called true bacteria. Eubacteria are usually found
in soil, water, living in and on of large organisms. Eubacteria are divided
into two groups known as gram positive and gram negative bacteria.
Archaebacteria are found in salt brines, ocean depths and hot springs. Three types of
archaebacteria are found: methanogens, halophiles and thermoacidophiles.
|
ArchaeaBacteria |
EuBacteria |
|
-Ancient bacteria- |
-True bacteria- |
Complexity |
Simple in their organization |
Complex than archaebacteria |
Habitat |
Can
sustain in extremely harsh environment such as oceans, hot springs,
marshlands, hot springs and gut of animals |
Found everywhere
- soil, organic matter, earth’s crust, water, bodies of animals and plants,
radioactive wastes, hot springs |
Size |
0.1-15 μm in diameter |
0.5-5 μm in diameter |
Shape |
spheres, rods, plates, spiral, flat or
square-shaped |
cocci, bacilli, vibrio, rods, filaments or spiral
in shape |
Cell wall |
Pseudopeptidoglycan |
Lipopolysaccharide/ Peptidoglycan with muramic
acid |
Membrane lipids |
Ether-linked, branched, aliphatic chains,
containing D-glycerol phosphate |
Ester-linked, straight chains of fatty acids,
containing L-glycerol phosphates |
RNA |
Consists
of single RNA |
Three types
of RNA |
RNA polymerase |
Complex subunit pattern |
Simple subunit pattern |
Introns (a long stretch of noncoding DNA found between exons (or coding regions) in a gene) |
Present in archaebacteria |
Absent in eubacteria |
Metabolism |
Methanogenesis- exhibit neither
glycolysis nor Kreb’s cycle |
Autotrophy, Aerobic and Anaerobic Respiration,
Fermentation and Photosynthesis-exhibit both glycolysis and Kreb’s cycle |
Reproduction and Growth |
Asexual
Reproduction, by fragmentation, budding and binary fission |
Other than binary fission, budding and
fragmentation, eubacteria can produce spores in order to remain dormant
during unfavorable conditions |
Types |
Methanogens, halophiles and thermophiles |
Gram positive and Gram negative |
Examples |
Halobacterium, Thermoproteus, Pyrobaculum,
Thermoplasma and Ferroplasma |
Mycobacteria, Bacillus, E. coli,
Pseudomonas, Clostridium etc |
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